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Tokyo-3 is being attacked by the third Angel. Conventional weapons prove ineffective, largely due to its projected force field called an AT Field. Nerv takes command of the battles, and is able to intercept and defeat the Angels using the Evangelions (Evas), biomechanical mecha previously developed in secret by Gehirn inside the underground Geofront; the Geofront is located underground and underneath Tokyo-3.
Not knowing why his father summoned him, Shinji Ikari, a 14 year old boy arrives to Tokyo-3 just as the Third Angel attacks the city. Shinji reluctantly agrees to join Nerv to pilot Evangelion Unit 01, and begins living with Captain Misato Katsuragi. He and Rei Ayanami battle the successive advances of the Angels together and are later joined by Asuka Langley Soryu, the pilot of Unit 02.
Each Eva has its own designated pilot (Unit 00 – Rei, Unit 01–Shinji, Unit 02–Asuka, and subsequently Unit 03–Toji Suzuhara), and operates by synchronizing the pilot's soul and the human soul inside the Eva via the enigmatic liquid substance known as LCL. (In the context of Evangelion, a "soul" refers to an individual's conscious existence, mental structure and identity, rather than a more conventional "supernatural" entity.) Surrounded by LCL, the pilot's nervous system, mind and body join with the Eva's controls, allowing the Eva to be controlled by the pilot's thoughts and actions. The higher a pilot's synchronization ratio, the better the pilot can control the Eva and fight more adeptly.
While Ritsuko mentions at the series' beginning that the Evas do have some biological components to them, the extent of this is not immediately apparent. Unit 01 is connected to Yui Ikari, Gendo's wife and Shinji's mother, since it absorbed her body and soul in a failed experiment, as shown in episodes 16 and 20. Rei herself is suspected to be a partial clone of Yui, and is known to harbor the soul of Lilith, the second Angel.
The Eva Unit 02 (piloted by Asuka Langley Soryu, accompanied by Shinji Ikari inside the plug) landing on the missile destroyer USS Ramage, while fighting the sixth Angel Gaghiel at sea.
It is finally revealed, towards the end of the series, that the Evas are not really "robots" but are actually cloned Angels (Units 00, 02, 03, and 04 are made from Adam, and 01 is made from Lilith) onto which mechanical components are incorporated as a means of restraint and control. This control is not perfect, as various units are shown over the course of the series driving into "berserker" mode, in which they can act of their own will, independent of any artificial power input.